Rather, odd-toed ungulates are known to rest all their body weight in three toes, or just the center toe. Even-toed ungulates on the other hand, rest their weight on their third and fourth toes.

They went extinct two million years ago and were part of the odd-toed ungulates, related to modern rhinoceroses, horses, and tapirs. The site uncovered fossil teeth and finger bones of the genus ...

Traditionally ungulates were considered even or odd-toed hoofed animals, however, as genetics demonstrated new relationships between species, the number of animals who qualify as ungulates expanded.